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Volume 8 Issue 1
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October 2004
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How do we see color?Electromagnetic radiation, varying in wavelength from gamma rays to microwaves, is constantly bombarding us from all directions. Our eyes are able to detect how much radiation is entering them, and from what direction, only if that radiation is within the visible spectrum, which is between approximately 380 and 780 nanometers (nm). The spectral power distribution (SPD) of a light source is graphical or tabulated data representing the amount of radiation emitted by a light source at each wavelength in the visible spectrum only. For example, Figure 2 shows the SPD of an incandescent lamp. The SPD provides the basic physical data needed to calculate light source color.
When people speak of color, they are usually discussing the appearance of an object (red, pink, purple, brown, white) or a light source (e.g., warm-white, cool-white). Although color appearance seems to come from the physical characteristics of the electromagnetic radiation reaching the retina, it is actually the result of signal processing performed by the visual system. More specifically, color appearance is the result of calculations performed by three separate "color channels" in the visual system. Each channel takes the same radiant power falling on the retina and processes it slightly differently. Research has produced a general model of the visual system, which provides a basic explanation of color appearance (CIE 2004). For light to stimulate the brain it must be absorbed by photoreceptors in the eye's retina. There are three types of cone photoreceptors responsible for color vision, each defined in large part by the photopigment contained within that photoreceptor. Figure 3 shows the spectral sensitivity of the three cone photoreceptors, L, M, and S. The letters symbolize the photoreceptors' peak spectral sensitivities to Long, Medium and Short wavelength radiation, respectively.
The neural signals generated by these photoreceptors are combined in different ways by the three color channels. One channel, the achromatic or luminance channel, calculates the amount of light falling on the retina from the sum of the outputs of the L and M cones. This luminance channel is related to the perceived brightness of the object or the light source. The other two channels, known as the color opponent channels, calculate the hues. One opponent channel subtracts the response of the L cones from that of the M cones to produce a Red versus Green response. When the signal strength of the channel is, for example, greater than zero, the channel signals "Red" to the brain; when the response is less than zero, it signals "Green" to the brain. The other opponent channel subtracts the response of the S cone from the sum of the L and M cones to produce a Blue versus Yellow response. All colors are seen by the brain as the following combinations: red or green; and blue or yellow. For example, orange is seen as reddish-yellow, while turquoise is seen as greenish-blue. Since both reds and greens are formed by only one color opponent channel, we can never see reddish-greens. Similarly, we can never see bluish-yellows. The color "white" is seen when both color opponent channels are balanced (i.e., the light is neither red or green, nor blue or yellow), so only the achromatic channel is signaling the brain. Although this model forms the foundation of color appearance, the actual appearance of a given color is based upon a much more complicated set of subtle interactions in the visual system. For example, a light may appear white when briefly flashed, but yellow when steadily viewed. A more common example of these neural interactions is the change in appearance of the same light at different adaptation levels. A light that looks orange at high light levels may look amber or brown under low light levels. A vivid example of how color appearance can change as a result of other neural activities in the brain is illustrated in Figure 4. The same physical stimuli (the colored squares) appear to be different depending upon the color of their surrounding.
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